Drag coefficient, but it doesn't operate on its own. The actual amount of drag is the coefficient relative to its frontal area. The frontal area is not reflected at all in the coefficient, and obviously isn't part of the marketing - because it is not particularly great on the Air. While maintaining the same coefficient, they could have designed dramatically different frontal areas (among other factors) to optimize for specific velocities. There are likely some other factors in downforce, tire sizes, etc they could tweak as well. These are non-trivial in impact at different velocities as well. These sort of things can dramatically impact rolling resistance at different velocities.
I do not know the details of what they chose to do of course, but it wasn't an accident that it ends up optimal at 70mph. Though maybe it's not even 70mph, but I'm pretty sure if they wanted to, they could tell you exactly what speed they designed for.
They almost certainly could change aspects of the car to make it more efficient at 80mph instead. It would likely then make it less efficient at other velocities.
You are correct that drag is the important factor, but the coefficient of drag is not the drag force, it is just a factor of it.